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Valley of the Giants is about the struggle between the evil Pennington (Charles Sellon), owner of a local railroad and Bruce Cardigan (Milton Sills), whose family owns a lumber mill. Pennington wants control of the Cardigan possessions, so he cuts off service to the mill. The Cardigans also need to obtain a franchise from the city council, which, too, is under the control of Pennington. This melodrama isn't always as heavy as it sounds. There are moments of humor, such as those offered by Arthur Stone who poses as a Chicago millionaire to help the Cardigans get their franchise. Sills' real-life wife, Doris Kenyon, plays the film's love interest, Shirley Pennington.