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Even though films up until the late '20s were silent, it doesn't mean they were devoid of musical devices. This sentimental drama uses a popular and poignant piece of music as its title and inspiration. You can bet the song was featured prominently in the musical score when the film played in theaters. Arnold Grahme (Dave Powell) is an organist who returns from a trip to discover that his sweetheart, Madeline (Alice Lake), has wed Count Zara (Henry Sedley). The marriage is not a happy one -- the count is a faithless brute who is carrying on an affair with his cousin, Pauline (Dagmar Godowsky). Grahme and the count wind up fighting a duel in Italy and the count is killed. Pauline spirits Madeline's daughter away, while Madeline enters a convent. Through his music, Grahme convinces her to marry him, but she dies shortly afterwards. Years later, Grahme's nephew, Jack Brown (Charles Mack), becomes involved with Joan, a chorus girl (Faire Binney). The couple argues and Jack leaves. Grahme falls in love with Joan, and, after finding out that she is Madeline's daughter, he proposes. When Jack returns, he can see that the young couple are really in love and he sacrifices his own happiness so that they can be together. While playing the organ, Madeline comes to him in spirit and tells him that through this sacrifice he has found the lost chord.